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it's all about different wheel standards...rather than gauge........

this is a problem that is still with us today, on modern stock........and is why there are a multitude of standards for each and every scale/gauge.

hence for 4mm scale, we have OO, OO finescale, EM, P4/S4...etc

even for HO we have proto 87 as well.....

3 mm scale has TT, plus 12mm gauge, 14mm gauge,e tc....

all differing standards within the same scale, as folk sought finer ,more realistic-looking wheels and track...and improved running of course.
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